English biologist Thomas Huxley (1825–1895) was a staunch supporter of Charles Darwin’s work; in fact, Huxley wrote a favorable review of Darwin’s On the Origin of Species that appeared soon after its publication. When the firestorm of controversy began after the appearance of Darwin’s book, Huxley was ready and able to defend Darwin, whose chronic public reticence about his theories was at that time exacerbated by illness. In 1860 at the British Association for the Advancement of Science, Huxley’s defense of Darwin was so vigorous during a debate with English Bishop Samuel Wilberforce (1805–1873) that he earned the title “Darwin’s bulldog.”